Publications

Names in bold indicate lab members; underlined names were undergraduate project students.

Tom’s Google Scholar profile is here.

2024

• Wilson, J.C., Wood, K.A., Griffin, L.R., Brides, K., Rees, E.C. and Ezard, T.H.G. (in press) Using satellite tracking to assess the use of protected areas and alternative roosts by Whooper and Bewick’s Swans. Ibis
• Brombacher, A., Poole, C.R., Ezard, T.H.G. & Wade, B.S. (in press) Heterochrony in the evolution of the planktonic foraminifera Trilobatus sacculifer plexus. Paleobiology.
• Lin, H., Zhang, W., Mulqueeney, J.M., Brombacher, A., Searle-Barnes, A., Nixon, M.S., Cai, X. and Ezard, T.H.G. (online) 3DKMI: A MATLAB Package to Generate Shape Signatures from Krawtchouk Moments. Methods. Ecol. Evol.
• Stott, I, Salguero-Gomez, R., Jones, O.R., Ezard, T.H.G., Gamelon, M., Lachish, S., Lebreton, J.-D., Simmonds, E.G., Gaillard, J.-M. & Hodgson, D.J. (2024) Life histories not just fast or slow. Trends Ecol. Evol..
• Mulqueeney, J., Searle-Barnes, A., Brombacher, A., Sweeney, M., Goswami, A. & Ezard T.H.G. (2024) How many specimens make a sufficient training set for automated 3D feature extraction? Open Science 11. doi: 10.1098/rsos.240113
• Jones, O.R., Ezard, T.H.G., Dooley. C.A., Healy, K., Hodgson, D.J., Mueller, M., Townley, S. & Salguero-Gomez, R. (2024) My family and other animals: Human demography under a comparative cross-species lens. In Human Evolutionary Demography edited by Oskar Burger, Ron Lee and Rebecca Sear. Published by the Center for Open Science. pp. 211-232.

2023

• Fabbrini, A., Greco, M., Iacoviello, F., Kucera, M., Ezard, T.H.G. & Wade, B.S. (2023) Bridging the extant and fossil record of planktonic foraminifera: implications for the Globigerina lineage. Paleontology e12676. doi: 10.1111/pala.12676.
• Avrutin, S., Goodwin, P.A. & Ezard, T.H.G. (2023) Assessing the Remaining Carbon Budget through the Lens of Policy-Driven Acidification and Temperature Targets. Climatic Change, 176, 128.
• Nicol-Harper, A., Doncaster, C.P., Wood, K., Hilton, G. & Ezard, T.H.G. (2023) Conservation implications of a mismatch between data availability and demographic impact. Ecol. & Evol.. doi: 10.1002/ece3.10269.
• Gascoigne, S.J.L., 38 others including Ezard, T.H.G. & Salguero-Gomez, R. (online) A standard protocol to report discrete stage-structured demographic information. Methods. Ecol. Evol. doi: 10.1111/2041-210X.14164.
• Searle-Barnes, A, Milton, J.A., Standish, C.D., Foster, G.L. & Ezard, T.H.G. (2023) Laser ablation mass spectrometry blast through detection in R. Rapid Comms. Mass. Spec. 37, e9533. doi: 10.1002/rcm.9533. R package.
• Yuen, S., Ezard, T.H.G. & Sobey, A.J. (2023) Epigenetic opportunities for evolutionary computation . Roy. Soc. Open Sci. 10, doi: 10.1098/rsos.221256.
• Brombacher, A., Wilson, P.A., Bailey, I. & Ezard, T.H.G. (2023) Morphological variation across space does not predict phenotypic change through time in two Neogene planktonic foraminifera species. Frontiers Ecol. Evol. 11, 1165174. doi: 10.3389/fevo.2023.1165174.
• Kearns, L.E., Searle-Barnes, A, Foster, G.L., Milton, J.A., Standish, C.D., & Ezard, T.H.G. (2023) The influence of geochemical variation among Globigerinoides ruber individuals on paleoceanographic reconstructions. Paleoceanogr. Paleoclimatol. 38, e2022PA004549. doi: 10.1029/2022PA004549.
• Saban, J, Romero, A, Ezard, T.H.G. & Chapman, M. (2023) Extensive crop-wild hybridisation during Brassica evolution, and selection during the domestication and diversification of Brassica crops. Genetics 223, iyad027. doi: 10.1093/genetics/iyad027.
• Brombacher, J.F.A., Searle-Barnes, A., Zhang, W., & Ezard, T.H.G. (2023) Analysing planktonic foraminiferal growth in three dimensions with foram3D: an R package for automated trait measurements from CT scans. J. Micropal. 41, 149.
• Kearns, L.E., Bohaty, S.M., Edgar, K. & Ezard, T.H.G. (2023) Small but mighty: how overlooked small species maintain community structure through middle Eocene climate change. Paleobiology 49, 77-98. doi: 10.1017/pab.2022.24

2022

• Brombacher, J.F.A., Schmidt, D. & Ezard, T.H.G. (2022) Developmental plasticity in deep time: a window to population ecological inference. Paleobiology 49, 259-270. doi: 10.1017/pab.2022.26
• Bataillon, T., Ezard, T.H.G., Kopp, M. & Masel, J. (2022) Genetics of adaptation & fitness landscapes: from toy models to testable quantitative predictions. Evolution 76, 1104-1107. doi: 10.1111/evo.14477

2021

• Nicol-Harper, A., Wood, K., Diamond, T., Major, H., Petersen, A., Tertitski, G., Doncaster, C.P., Ezard, T.H.G., Hilton, G. (2021). Vital rate estimates for the common eider Somateria mollissima, a data-rich exemplar of the seaduck tribe. Ecol Solut Evidence 2, e12108.
• Fabbrini, A., Zaminga, I., Ezard, T.H.G. & Wade, B.S. (2021) Systematic Taxonomy of middle Miocene Sphaeroidinellopsis (planktonic foraminifera). J Syst Palaeont 19, 953-968.
• Kearns, L.E., Bohaty, S.M., Edgar, K., Nogue, S., Ezard, T.H.G. (2021). Searching for function: Reconstructing adaptive niche changes using geochemical and morphological data in planktonic Foraminifera. Front. Ecol. Evol.. doi: 10.3389/fevo.2021.679722
• Brombacher, J.F.A., Wilson, P.A., Bailey, I. & Ezard, T.H.G. (2021). The Dynamics of Diachronous Extinction Associated With Climatic Deterioration Near the Neogene/Quaternary Boundary. Paleoceanography and Paleoclimatology, 36, e2020PA004205

2020

• Jakob, K.A., Wilson, P.A., Pross, J., Ezard, T.H.G., Fiebig, J., Repschläger, J. & Friedrich, O. (2020) A new sea-level record for the Neogene/Quaternary boundary reveals transition to a more stable East Antarctic Ice Sheet. Proc Natl Acad Sci USA 117, 30980.
• Rillo, M.C., Miller, C.G., Kucera, M. & Ezard, T.H.G. (2020) Intraspecific size variation in planktonic foraminifera cannot be consistently predicted by the environment. Ecol. & Evol. 10, 11579.
• Ezard, T.H.G. (2020) Book Review: Biodemography: An introduction to concepts and methods by Carey & Roach. Biom. J. 62, 1359.
• Zhang, W., Ezard, T.H.G., Searle-Barnes, A., Brombacher, A., Katsamenis, O. & Nixon, M.S. (2020) Towards Understanding Speciation By Automated Extraction And Description Of 3d Foraminifera Stacks. 2020 IEEE Southwest Symposium on Image Analysis and Interpretation, 30.
• Che-Castaldo, J., Jones, O.R., 13 others including Ezard, T.H.G. & Salguero-Gómez, R. (2020) Comments to “Persistent problems in the construction of matrix population models”. Ecol. Model. 416, 108913.

2019

• Rillo, M.C., Sugawara, M.T.C., Cabella, B., Jonkers, L., Baranowski, U.K., Kucera M, & Ezard, T.H.G.. (2019). On the mismatch in the strength of competition among fossil and modern species of planktonic foraminifera. Global Ecol. Biogeog. 28, 1866.
• Healy, K., Ezard, T.H.G., Jones, O.R., Salguero-Gómez, R. & Buckley, Y.M. (2019) Animal life history is shaped by the pace of life and distribution of age-specific mortality and reproduction Nature Ecol. Evol. 3, 1217.
• Rillo, M.C., Kucera, M., Ezard, T.H.G. & Miller, C.G. (2019) Surface Sediment Samples From Early Age of Seafloor Exploration Can Provide a Late 19th Century Baseline of the Marine Environment Frontiers Marine Sci 5, 517.

2018

• Brombacher, A., Wilson, P.A., Bailey, I. & Ezard, T.H.G. (2018) Temperature is a poor proxy for synergistic climate forcing of plankton evolution. Proc. Roy. Soc. B 285, 20180665.
• Brombacher, A., Elder, L.E., Hull, P.M., Wilson, P.A. & Ezard, T.H.G. (2018) Calibration of test diameter and area as proxies for body size in the planktonic foraminifer Globoconella puncticulata. J. Foram. Res. 48, 241.
• Nicol-Harper A., Dooley C.A., Packman D., Mueller, M., Bijak, J., Hodgson, D.J., Townley S. & Ezard T.H.G. (2018) Inferring transient dynamics of human populations from non-normality metrics. Popul. Ecol. 60, 1.

2017

• Edgar, K.M., Hull, P.M. & Ezard, T.H.G. (2017) Evolutionary history biases inferences of ecology and environment from δ13C but not δ18O values. Nature Comms 8, 1106. R code and data compilations on FigShare.
• Pigeon, G., Ezard, T.H.G., Festa-Bianchet, M., Coltman, D.W., & Pelletier, F. (2017) Fluctuating effects of genetic and plastic changes in body mass on population dynamics in a large herbivore. Ecology 98, 2456-2467.
• Brombacher, A., Wilson, P.A., Bailey, I. & Ezard, T.H.G. (2017) The breakdown of static and evolutionary allometries during climate upheaval. Amer Nat 190, 350. Data available on Dryad. Charles Downie Award 2017 for “best & most significant” micropalaeontogical paper.
• Brombacher, A., Wilson, P.A. & Ezard, T.H.G. (2017) Calibration of the repeatability of foraminiferal test size and shape measures with recommendations for future use. Mar. Micropal. 133, 21.
• McDonald, J.L., Franco, M., Townley, S., Ezard, T.H.G., Jelbert, K. & Hodgson, D.J. (2017) Divergent demographic strategies of plants in variable environments. Nature Ecol. & Evol. 1, 0029.
• Rillo, M.C., Whittaker, J., Ezard, T.H.G., Purvis, A., Henderson, A. S., Stukins, S. & Miller, C.G. (2017). The unknown planktonic foraminiferal pioneer Henry A. Buckley and his collection at The Natural History Museum, London. J. Foram. Res. 36, 191.

2016

• Cooke, R.S.C., Woodfine, T., Petretto, M. & Ezard, T.H.G. (2016) Resource partitioning between ungulate populations in arid environments. Ecol. Evol. 6, 6354.
• Ezard, T.H.G. & Purvis, A. (2016) Environmental changes define ecological limits to species richness and the mode of macroevolutionary competition. Ecol. Lett.. 19, 899. R code and data accessibility.
• Ezard, T.H.G., Quental, T.B., & Benton, M.B. (2016) The challenges to inferring the regulators of biodiversity in deep time. Phil. Trans. Roy. Soc. B. 371, 20150216. Introduction to our Theme Issue on The Regulators of Biodiversity in Deep Time.

2015

• Spake, R., Ezard, T.H.G., Martin, P.A., Newton, A.C. & Doncaster, C.P. (2015) A meta-analysis of functional group responses to forest recovery outside of the tropics. Cons. Biol. 29, 1695.
• Ezard, T.H.G., Edgar, K.M. & Hull, P.M. (2015) Environmental and biological controls on size-specific δ13C and δ18O in recent planktonic foraminifera. Paleooceanography 30, 151. R code and data compilations on FigShare.
• Jørgensen, P.S. and 14 others including Ezard T.H.G. (2015). Connecting people and ideas from around the world: Global innovation platforms for next-generation ecology and beyond. Ecosphere 6:art68.

2014

• Prizak, R., Ezard, T.H.G. & Hoyle, R.B. (2014). Fitness consequences of maternal and grandmaternal effects. Ecol. Evol. 4, 3139. MATLAB simulation routines on figshare.
• Barraquand, F. Ezard, T.H.G., Joergensen, P., Zimmerman, N., Chamberlain, S.A., Salguero-Gómez, R., Curran, T.J. & Poisot, T. (2014) Lack of quantitative training among early-career ecologists: a survey of the problem and potential solutions. PeerJ 2:e285.
• Ezard, T.H.G., Prizak, R. & Hoyle, R.B. (2014). The fitness implications of adaptation via phenotypic plasticity and maternal effects. Func. Ecol. 28, 693. MATLAB simulation routines on figshare.
• Seddon, A.W.R., Mackay, A.W., Baker, A.G., 32 others, then Ezard, T.H.G. & then another 33 others (2014) Looking forward through the past: Identification of fifty priority research questions in palaeoecology. J. Ecol.. 102, 256.
• Pearson, P.N. & Ezard, T.H.G. (2014) Evolution and speciation in the Eocene planktonic foraminifer Turborotalia. Paleobiology. 40, 130. Supporting data, movies & figures on figshare.

2012-2013

• Ezard, T.H.G., Thomas, G.H. & Purvis, A. (2013) Inclusion of a near-complete fossil record reveals speciation-related molecular evolution. Methods Ecol. Evol. 4, 745. Phylogenies & data via figshare. Daily Mail and Guardian Higher Education Network press coverage].
• Aze, T., Ezard, T.H.G., Purvis, A., Coxall, H.K., Stewart, D.R.M., Wade, B.S. & Pearson, P.N. (2013). Identifying anagenesis and cladogenesis in the fossil record. Proc. Natl. Acad. Sci USA 110, E2946.
• Townley, S. & Ezard, T.H.G. (2013). A G matrix analogue to capture the cumulative effects of non-genetic inheritance. J. Evol. Biol. 26, 1234.
• Hoyle, R.B. & Ezard, T.H.G. (2012). The benefits of maternal effects in novel and in stable environments. J. Roy. Soc. Interface 9, 2403. Podcast for the Journal of Ecology. MATLAB simulation routines on figshare.
• Ezard, T.H.G., Pearson, P.N., Aze, T., & Purvis, A. (2012). The Meaning of Birth and Death (in Macroevolutionary Birth-Death Models). Biol. Lett. 8, 139.

2010-2011

• Ezard, T.H.G., Aze, T., Pearson, P.N. & Purvis, A. (2011). Interplay between Changing Climate and Species’ Ecology Drives Macroevolutionary Dynamics. Science 332, 349. Faculty of 1000 “must-read”. Pecha Kucha talk.
• Aze, T., Ezard, T.H.G., Purvis, A., Coxall, H.K., Stewart, D.R.M., Wade, B.S. & Pearson, P.N. (2011). A phylogeny of Cenozoic macroperforate planktonic foraminifera from fossil data. Biol. Rev. 86, 900. Charles Downie Award 2011 for “best & most significant” micropalaeontogical paper.
• Ezard, T.H.G., Pearson, P.N. & Purvis, A. (2010). Algorithmic approaches to aid species delimitation in multidimensional morphospace. BMC Evol. Biol., 10: 175.
• Ezard, T.H.G., Bullock, J.M., Dalgleish, H.J., Millon, A., Pelletier, F., Ozgul, A. & Koons, D.N. (2010). Matrix models for a changeable world: the importance of transient dynamics in population management. J. Appl. Ecol. 47, 515.
• Ezard, T.H.G. & Coulson, T. (2010). How sensitive are elasticities of long-run stochastic growth to how environmental variability is modelled? Ecol. Model. 221, 191.

2008-2009

• Ezard, T.H.G. & Purvis, A. (2009) paleoPhylo: free software to draw paleobiological phylogenies. Paleobiology 35, 460.
• Ezard, T.H.G., Côté, S.D. & Pelletier, F. (2009). Eco-evolutionary dynamics: disentangling phenotypic, environmental and population fluctuations. Phil. Trans. R. Soc. Lond. B 364, 1491.
• Ezard, T.H.G., Gaillard, J.-M., Crawley, M.J. & Coulson, T. (2008) Habitat dependence and correlations between elasticities of long-term growth rates. Am. Nat. 172: 424. Bright Club Guildford reworking as a stand-up comedy set.
• Coulson, T., Ezard, T.H.G., Pelletier, F., Stenseth, N.C., Pilkington, J. Pemberton, J.M., Kruuk, L.E.B., Clutton-Brock, T.H. & Crawley, M. (2008) Estimating the functional form for the density dependence from life history data. Ecology 89: 1664.
• Becker, P.H., Ezard, T.H.G., Ludwigs, J.-D., Sauer-Gürth, H. & Wink, M. (2008) Population sex ratio shift from fledging to recruitment: consequences for demography in a philopatric seabird. Oikos 117: 60.

2006-2007

• Ezard, T.H.G., Becker, P.H. & Coulson, T. (2007) Correlations between Age, Phenotypic Traits and Individual Contributions to Population Growth in Common Terns. Ecology 88: 2496.
• Dittmann, T., Ezard, T.H.G. & Becker, P.H. (2007) Prospectors colony attendance is sex-specific and increases future recruitment chances in a seabird. Behav. Proc. 76: 198
• Ezard, T.H.G., Becker, P.H. & Coulson, T. (2006). The Contributions of Age and Sex to Variation in Common Tern Population Growth Rate. J. Anim. Ecol. 75: 1379.
• Ezard, T.H.G. & Travis, J.M.J. (2006). The Impact of Habitat Loss and Fragmentation on Genetic Drift and Fixation Time. Oikos 114: 376.
• Travis, J.M.J. & Ezard, T.H.G., (2006). Habitat geometry, population viscosity and the rate of genetic drift. Ecol. Inf. 1, 153.